The suggested citation for this work is listed below:

Mills, C.E. Internet 1999-present. Stauromedusae: list of all valid species names. Electronic internet document available at http://faculty.washington.edu/cemills/Staurolist.html. Published by the author, web page established October 1999, last updated (see date at end of page).

Having put together a list of what I believe to be all valid scientific names for the Ctenophora, I decided to compile a similar list of names for the Stauromedusae (Phylum Cnidaria: Staurozoa) that seem to be valid at the present time; links are included to stauromedusa images located elsewhere on the web. Whereas I believe that the list of genera and species is fairly complete, placement of these genera into families and subfamilies seems still to be fairly plastic, with need for further revision indicated. The basic structure of this taxonomy comes from Oskar Carlgren (1935) and Tohru Uchida (1929). Few living authors (including myself) have seen many species of Stauromedusae, which has been one problem in interpreting relationships. Presenting here an Internet webpage rather than a scientific paper, I have tried to be consistent in interpreting previous authors' work at the family and subfamily levels. Dr. Yayoi Hirano of Japan, perhaps the best-informed stauromedusan biologist in the world at this point, has offered some comments and corrections to this list.

There are about 50 known species of stauromedusae, which range from the intertidal zone to deep hydrothermal vent habitats. In some cases here, I have listed synonyms to good species names in parentheses on the line following - the list of synonyms may not be complete.

Note that genus and species names are always written in italics. If the author and date following a species name are within parentheses, that is the taxonomists' code for indicating that the species has been moved from the genus in which it was originally described; author and date not in parentheses indicates that the genus has not changed from the original description. (I am not making this format up - it is strictly formalized and governed by an international body of taxonomists.)

If you use this list in your studies, I would appreciate it if you cite it as suggested above as an electronic publication. The list is an ongoing work and is subject to modification at any time. I consider it my intellectual property and do not want to find it dumped wholesale onto someone else's web page - links to it are fine. I welcome any additions or corrections that anyone may wish to make - please contact me at the above address. In order to be true to its in-progress nature, I have left my personal shorthand in the electronic manuscript, so any entry preceeded by . means that I (or Yayoi Hirano) have not yet been able to check the original citation for accuracy. Although I have chosen to put this list of species names and authors on the web, it will be noted that I have not included the bibliography from which it derives.

I have included links to the few images of stauromedusae located elsewhere on the web. Any additions to these links are welcome.

Phylum CNIDARIA

newest classification = Class Staurozoa Marques and Collins, 2004

older classification = Class Scyphozoa

. Scyphomedusae Lankaster, 1877

. Order Stauromedusae Haeckel, 1879

(=Lucernariae Clark, 1863?)

Suborder Eleutherocarpida Clark, 1863

. Family Lucernariidae Johnston, 1847

Subfamily Lucernariinae Carlgren, 1935

Haliclystus Clark, 1863

antarcticus Pfeffer, 1889

auricula (Rathke, 1806) (another image - drawing)

borealis Uchida, 1933

kerguelensis Vanhöffen, 1908

monstrosus (Naumov, 1961)

(=Octomanus Naumov, 1961)

. octoradiatus (Lamarck, 1816)

salpinx Clark, 1863

"sanjuanensis" Hyman, 1940 or Hirano, 1997 or undescribed?

sinensis Ling 1937

stejnegeri Kishinouye, 1899

tenuis Kishinouye, 1910

sp. ("californensis" of Gwilliam, 1956, unpublished)

sp. undescribed, referred to by Hirano, 1997

Lucernaria O. F. Müller, 1776

australis Vanhöffen, 1908

. bathyphila Haeckel, 1879

haeckeli (Antipa, 1891)

. infundibulum Haeckel, 1880

janetae Collins and Daly, 2005 (another image)

quadricornis O. F. Müller, 1776 this image is probably L. quadricornis

(?=L. pyramidalis Haeckel)

. saint-hilarei (Radicorzew)

walteri (Antipa, 1891)

(includes kükenthali (Antipa, 1891) - synonymized by Kramp, 1961)

Stenoscyphus Kishinouye, 1902

inabai (Kishinouye, 1893)

Stylocoronella Salvini-Plawen, 1966

riedli Salvini-Plawen, 1966

variabilis Salvini-Plawen, 1987

Undescribed subfamily "Stenoscyphopsinae" of Gwilliam, 1956, unpublished

Undescribed genus ("Stenoscyphopsis" of Gwilliam, 1956, unpublished)

Undescribed species ("vermiformis" of Gwilliam, 1956, unpublished)

Family Kishinouyeidae Uchida, 1929

or Subfamily Kishinouyeinae Carlgren, 1935

Kishinouyea Mayer, 1910

corbini Larson, 1980

hawaiiensis Edmondson, 1930

nagatensis (Oka, 1897)

Lucernariopsis Uchida, 1929

(=Calvadosia Clark, 1863)

. campanulata (Lamouroux, 1815)

(=Lucernaria discoidea Eales, 1938)

capensis Carlgren, 1938

cruxmelitensis Corbin, 1978

vanhoeffeni (Browne, 1910)

. sp. of Panikkar, 1944

Sasakiella Okubo, 1917

cruciformis Okubo, 1917

tsingtaoensis Ling, 1937

. Family Lipkeidae Vogt, 1887

or Subfamily Lipkeinae Carlgren, 1935

Lipkea Vogt, 1886

(=Capria Antipa, 1893)

ruspoliana Vogt, 1886

stephensoni Carlgren, 1933

sturdzii (Antipa, 1893)

Family Kyopodiidae Larson, 1988

Kyopoda Larson, 1988

lamberti Larson, 1988

Suborder Cleistocarpida Clark, 1863

Family Depastridae Haeckel, 1879

. Subfamily Depastrinae Uchida

Depastromorpha Carlgren, 1935

africana Carlgren, 1935

. Depastrum Gosse, 1858

. (=Depastrella ?) - see Carlgren, 1933

. cyathiforme (M. Sars, 1846)

Subfamily Thaumatoscyphinae Carlgren, 1935

Manania Clark, 1863

(=Brochiella Krumbach, 1925)

(=Thaumatoscyphus, Kishinouye, 1910)

atlantica (Berrill, 1962)

auricula (Fabricius, 1780)

(=Halimocyathus lagena (O. F. Müller, 1776))

distincta (Kishinouye, 1910)

(=Thaumatoscyphus uchidai Naumov, 1961) by Hirano, 1986

gwilliami Larson and Fautin, 1989 (bottom image)

handi Larson and Fautin, 1989 (another photo=top image, red color instead of green)

hexaradiata (Broch, 1907)

. lagena (?) - see Carlgren, 1933

(=Halimocyathus lagena author?) - see Carlgren, 1933

uchidai Naumov, 1961 (Hirano now believes this is a distinct species)

Halimocyathus Clark, 1863

platypus Clark, 1863

Subfamily Craterolophinae Uchida, 1929 or Family Craterolophidae

Craterolophus Clark, 1863

. convolvulus (Johnston, 1835) (another image - drawing)

(=C. tethys Clark, 1863)

macrocystis von Lendenfeld, 1884