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Campos, E. Octavio, Bradshaw, H.D., Jr., & Daniel, Thomas L. (2015) Shape matters: Corolla curvature improves nectar discovery in the hawkmoth (Manduca sexta).  Functional Ecology 29: 462-468.

Byers, Kelsey J.R.P., Vela, James P., Peng, Foen, Riffell, Jeffrey A., & Bradshaw, H.D., Jr. (2014) Floral volatile alleles can contribute to pollinator-mediated reproductive isolation in monkeyflowers (Mimulus).  The Plant Journal 80: 1031-1042.

Yuan, Yao-Wu, Sagawa, Janelle M., Frost, Laura, Vela, James P., & Bradshaw, H.D., Jr. (2014) Transcriptional control of floral anthocyanin pigmentation in monkeyflowers (Mimulus).  New Phytologist 204: 1013-1027.

Yuan, Yao-Wu, Byers, Kelsey J.R.P., & Bradshaw, H.D., Jr. (2013) The genetic control of plant-pollinator interactions.  Current Opinion in Plant Biology 16: 422-428.

Yuan, Yao-Wu, Sagawa, Janelle M., Di Stilio, Veronica S., & Bradshaw, H.D., Jr. (2013) Bulk segregant analysis of an induced floral mutant identifies a MIXTA-like R2R3 MYB controlling nectar guide formation in Mimulus lewisii.   Genetics 194: 523-528.

Yuan, Yao-Wu, Sagawa, Janelle M., Young, Riane C., Christensen, Brian J., & Bradshaw, H.D., Jr. (2013) Genetic dissection of a major anthocyanin QTL contributing to pollinator-mediated reproductive isolation between sister species of Mimulus.   Genetics 194: 255-263.

Owen, Christina R. & Bradshaw, H.D., Jr. (2011) Induced mutations affecting pollinator choice in Mimulus lewisii (Phrymaceae). Arthropod-Plant Interactions 5(3): 235-244.

Angert, A.L., Bradshaw, H.D., Jr., & Schemske, D.W. (2008) Using experimental evolution to investigate geographic range limits in monkeyflowers.  Evolution 62: 2660-2675.

Bradshaw, H.D., Jr. (2005) Mutations in CAX1 produce phenotypes characteristic of plants tolerant to serpentine soils.  New Phytologist 167: 81-88.

Brady, K.U, Kruckeberg, A.R., & Bradshaw, H.D., Jr. (2005) Evolutionary ecology of plant adaptation to serpentine soils.  Annual Review of Ecology, Evolution, and Systematics 36: 243-266.

Strauss, S.H. & Bradshaw, H.D., Jr., eds. (2004) The bioengineered forest: Challenges for science and society. Resources for the Future Press, Washington, D.C. ISBN 1-891853-71-6.

Bradshaw, H.D., Jr. and Schemske, D.W. (2003) Allele substitution at a flower colour locus produces a pollinator shift in monkeyflowers.  Nature 426: 176-178.

Ramsey, J. R., Bradshaw, H. D., Jr., & Schemske, D. W. (2003) Components of reproductive isolation in the monkeyflowers Mimulus lewisii and M. cardinalis (Phrymaceae).  Evolution 57(7): 1520-1534.

Stirling, B., Yang, Z.K., Gunter, L.E., Tuskan, G.A., & Bradshaw, H.D., Jr. (2003) Comparative sequence analysis between orthologous regions of the Arabidopsis and Populus genomes reveals substantial synteny and collinearity.  Canadian Journal of Forest Research 33(11): 2245-2251.

Ferris, R., Long, L., Bunn, S.M., Robinson, K.M., Bradshaw, H.D., & Taylor, G. (2002) Leaf stomatal and epidermal cell development: identification of putative quantitative trait loci in relation to elevated carbon dioxide concentration in poplar.  Tree Physiology 22: 633-640.

Bradshaw, H.D., Jr. and Strauss, S.H. (2001) Plotting a course for GM forestry.  Nature Biotechnology 19(12): 1103-1104.

Stirling, B., Newcombe, G., Vrebalov, J., Bosdet, I., & Bradshaw, H.D., Jr. (2001) Suppressed recombination around the MXC3 locus, a major gene for resistance to poplar leaf rust.  Theoretical and Applied Genetics 103(8): 1129-1137.

Newcombe, G., Stirling, B., and Bradshaw, H.D., Jr. (2001) Abundant pathogenic variation in the new hybrid rust population of Melampsora xcolumbiana on hybrid poplar.  Phytopathology 91: 981-985.

Bradshaw, H.D., Jr. and Schemske, D.W. (2001) The birds and the bees of plant evolution.  Northwest Science and Technology Spring 2001: 44-49.

Bradshaw, H.D., Jr., Ceulemans, R., Davis, J., and Stettler, R.F. (2000) Emerging model systems: Poplar (Populus) as a model forest tree.  Journal of Plant Growth Regulators 19(3): 306-313.

Frewen, B.E., Chen, T.H.H., Howe, G., Davis, J., Rohde, A., Boerjan, W., & Bradshaw, H.D., Jr. (2000) QTL and candidate gene mapping of bud set and bud flush in Populus. Genetics 154: 837-845.

Newcombe, G, Stirling, B., McDonald, S., and Bradshaw, H.D., Jr. (2000) Melampsora xcolumbiana, a natural hybrid of M. medusae and M. occidentalis.  Mycological Research 104(3): 261-274.

Schemske, D.W. & Bradshaw, H.D., Jr. (1999) Pollinator preference and the evolution of floral traits in monkeyflowers (Mimulus). Proceedings of the National Academy of Sciences USA 96(21): 11910-11915.

Howe, G.T., Davis, J., Jeknic, Z., Chen, T.H.H., Frewen, B., Bradshaw, H.D., Jr., & Saruul, P. (1999) Physiological and genetic approaches to studying endodormancy-related traits in Populus. HortScience 34(7): 1174-1184.

Bradshaw, H.D., Jr., Otto, K.G., Frewen, B.E., McKay, J.K. & Schemske, D.W. (1998) Quantitative trait loci affecting differences in floral morphology between two species of monkeyflower (Mimulus). Genetics 149: 367-382.

Bradshaw, H.D., Jr. (1998) Case history in genetics of long-lived plants: Molecular approaches to domestication of a fast-growing forest tree: Populus. In: Molecular dissection of complex traits. Paterson, A. (ed), CRC Press, Boca Raton, Florida, pp. 219-228.

Wu, R., Bradshaw, H.D., Jr., & Stettler, R.F. (1998) Developmental quantitative genetics of growth in Populus. Theoretical and Applied Genetics 97: 1110-1119.

Wilbert, S.M., Schemske, D.W., & Bradshaw, H.D., Jr. (1997) Floral anthocyanins from two monkeyflower species with different pollinators. Biochemical Systematics and Ecology 25(5): 437-443.

Wu, R., Bradshaw, H.D., Jr., & Stettler, R.F. (1997) Molecular genetics of growth and development in Populus. V. Mapping quantitative trait loci affecting leaf variation. American Journal of Botany 84(2): 143-153.

Newcombe, G. & Bradshaw, H.D., Jr. (1996) Quantitative trait loci conferring resistance in hybrid poplar to leaf spot caused by Septoria populicola. Canadian Journal of Forest Research 26(11): 1943-1950.

Bradshaw, H.D., Jr. (1996) Molecular genetics of Populus. In: Biology of Populus and its implications for management and conservation. Stettler, R.F., Bradshaw, H.D., Jr., Heilman, P.E., & Hinckley, T.M. (eds), NRC Press, Ottawa, pp. 183-199.

Villar, M., Lefevre, F., Bradshaw, H.D., & Teissier du Cros, E. (1996) Molecular genetics of rust resistance in poplars (Melampsora larici-populina Kleb./Populus sp.) by bulked segregant analysis in a 2 x 2 factorial mating design. Genetics 143: 531-536.

Newcombe, G., Bradshaw, H.D., Jr., Chastagner, G.A., & Stettler, R.F. (1996) A major gene for resistance to Melampsora medusae f.sp. deltoidae in a hybrid poplar pedigree. Phytopathology 86(1): 87-94.

Bradshaw, H.D., Jr., Wilbert, S.M., Otto, K.G., & Schemske, D.W. (1995) Genetic mapping of floral traits associated with reproductive isolation in monkeyflowers (Mimulus). Nature 376: 762-765.

Bradshaw, H.D., Jr. & Stettler, R.F. (1995) Molecular genetics of growth and development in Populus. IV. Mapping QTLs with large effects on growth, form, and phenology traits in a forest tree. Genetics 139: 963-973.

Bradshaw, H.D., Jr., Villar, M., Watson, B.D., Otto, K.G., Stewart, S., & Stettler, R.F. (1994) Molecular genetics of growth and development in Populus. III. A genetic linkage map of a hybrid poplar composed of RFLP, STS, and RAPD markers. Theoretical and Applied Genetics 89(5): 167-178.

Bradshaw, H.D., Jr. & Coulson, T.D. (1988) Male-biased sex ratio in captive-bred Harris' Hawks. Journal of Raptor Research 22: 116-117.

Bradshaw, H.D., Jr. (1983) Molecular cloning and cell cycle-specific regulation of a functional human thymidine kinase gene. Proceedings of the National Academy of Sciences USA 80: 5588-5591. 

Last updated on 4 November 2014.  Send questions or comments to