publications
 
  Halpern, C. B., J. A. Antos, D. McKenzie, and A. M. Olson. 2016. Past tree influence and prescribed fire mediate biotic interactions and community reassembly in a grassland-restoration experiment. Journal of Applied Ecology 53:264-273. (Abstract)  
  Baker, S. C., C. B. Halpern, T. J. Wardlaw, R. E. Bigley, R. L. Crawford, G. J. Edgar, S. A Evans, J. F. Franklin, G. J. Jordan, Y. Karpievitch, T. A. Spies, and R. J. Thompson. 2015. Short- and long-term benefits for biodiversity of retaining unlogged patches in harvested areas. Forest Ecology and Management 353:187-195. (Abstract)  
  Harmon, M. E., B. Fasth, C. B. Halpern, and J. A. Lutz. 2015. Uncertainty analysis: an evaluation metric for synthesis science. Ecosphere 6(4):63. (pdf)  
  Halpern, C. B., J. A. Antos, and L. M. Beckman. 2014. Vegetation recovery in slash-pile scars following conifer removal in a grassland-restoration experiment. Restoration Ecology 22:731-740. (pdf)  
  Halpern, C. B., M. Dovciak, L. S. Urgenson, and S. A. Evans. 2014. Substrates mediate responses of forest bryophytes to a gradient in overstory retention. Canadian Journal of Forest Research 44:855-866. (Abstract)  
  Urgenson, L. S., S. H. Reichard, and C. B. Halpern. 2014. Habitat factors and species’ traits influence riparian community recovery following removal of Bohemian knotweed (Polygonum x bohemicum). Northwest Science 88:246-260. (Abstract)  
  Kremer, N. J., C. B. Halpern, and J. A. Antos. 2014. Conifer reinvasion of montane meadows following experimental tree removal and prescribed burning. Forest Ecology and Management 319:128-137. (Abstract)  
  McCain, C., C. B. Halpern, and S. Lovtang. 2014. Non-forested plant communities of the northern Oregon Cascades. USDA Forest Service, Pacific Northwest Region, Technical Paper R6-ECOL-TP-01-14. (Link to pdf)  
  Case, M. F., C. B. Halpern, and S. A. Levin. 2013. Contributions of gopher mound and casting disturbances to plant community structure in a Cascade Range meadow complex. Botany 91:555-561. (Abstract)  
  Halpern, C. B., and J. A. Lutz. 2013. Canopy closure exerts weak controls on understory dynamics: A 30-year study of overstory-understory interactions. Ecological Monographs 83:221-237. (pdf)  
  Haugo, R. D., J. D. Bakker, and C. B. Halpern. 2013. Biotic interactions regulate conifer invasion of grasslands. Forest Ecology and Management 289:175-182. (Abstract)  
  Urgenson, L. S., C. B. Halpern, and P. D. Anderson. 2013. Level and pattern of overstory retention influence rates and forms of tree mortality in mature, coniferous forests of the Pacific Northwest, U.S.A. Forest Ecology and Management 308:116-27. (Abstract)  
  Urgenson, L. S., C. B. Halpern, and P. D. Anderson. 2013. Twelve-year responses of planted and naturally regenerating conifers to variable-retention harvest in the Pacific Northwest, U.S.A. Canadian Journal of Forest Research 43:46-55. (Abstract)  
  Halpern, C. B., J. Halaj, S. A. Evans, and M. Dovciak. 2012. Level and pattern of overstory retention interact to shape long- term responses of understories to timber harvest. Ecological Applications 22:2049-2064. (pdf)  
  Evans, S. A., C. B. Halpern, and D. McKenzie. 2012. The contributions of forest structure and substrate to bryophyte diversity and abundance in mature coniferous forests of the Pacific Northwest. The Bryologist 115:278-294. (Abstract)  
  Halpern, C. B., R. D. Haugo, J. A. Antos, S. S. Kaas, and A. L. Kilanowski. 2012. Grassland restoration with and without fire: evidence from a tree-removal experiment. Ecological Applications 22:425-441 (pdf) (Appendices)  
  Holloway, G. L., W. P. Smith, C. B. Halpern, R. A. Gitzen, C. C. Maguire, and S. D. West. 2012. Influence of forest structure and experimental green–tree retention on northern flying squirrel (Glaucomys sabrinus) abundance. Forest Ecology and Management 285:187-194. (Abstract)  
  Rice, J. M., C. B. Halpern, J. A. Antos, and J. A. Jones. 2012. Spatio-temporal patterns of tree establishment are indicative of  biotic interactions during early invasion of a montane meadow. Plant Ecology 213:555-568. (Abstract)  
  Urgenson. L. S., S. H. Reichard, and C. B. Halpern. 2012. Multiple competitive mechanisms underlie the effects of a strong invader on early- to late-seral tree seedlings. Journal of Ecology 100:1204-1215. (Abstract)  
  Bustamante-Sánchez, M. A., J. J. Armesto, and C. B. Halpern. 2011. Biotic and abiotic controls on tree colonization in three early successional communities of Chiloé Island, Chile. Journal of Ecology 99:288-299. (Abstract)  
  Haugo, R. D., C. B. Halpern, and J. D. Bakker. 2011. Landscape context and tree influences shape the long-term dynamics of forest-meadow ecotones in complex mountain ecosystems. Ecosphere 2(8):91.(pdf)  
  Dovciak, M., and C. B. Halpern. 2010. Diversity-stability relationships in forest herb populations during four decades of community assembly. Ecology Letters 13:1300-1309. (Abstract)  
  Halpern, C. B., J. A. Antos, J. M. Rice, R. D. Haugo, and N. L. Lang. 2010. Tree invasion of a montane meadow complex: temporal trends, spatial patterns, and biotic interactions. Journal of Vegetation Science 21:717-732. (Abstract)  
  Haugo, R. D., and C. B. Halpern. 2010. Tree age and tree species shape positive and negative interactions in a montane meadow. Botany 88:488-499. (Abstract)  
  Jones, C. C., S. A. Acker, and C. B. Halpern. 2010. Combining local- and large-scale models to predict the distributions of invasive plant species. Ecological Applications 20:311-326. (pdf)  
  Webster, K. A., and C. B. Halpern. 2010. Positive long-term responses of vegetation to reintroduction and repeated use of fire in mixed-conifer forests of the Sierra Nevada. Ecosphere 1(5):9. (pdf)  
 

Sprugel, D. G., K. G. Rascher, R. Gersonde, M. Dovciak, J. A. Lutz, and C. B. Halpern. 2009. Spatially explicit modeling of overstory manipulations in young forests: Effects on stand structure and light. Ecological Modelling 220:3565-3575. (Abstract)

 
  Aubry, K. B., C. B. Halpern, and C. E. Peterson. 2009. Variable-retention harvests in the Pacific Northwest: a review of short-term findings from the DEMO study. Forest Ecology and Management 258:398-408. (Abstract)  
  Compagnoni, A., and C. B. Halpern. 2009. Properties of native plant communities do not determine exotic success during early forest succession. Ecography 32:449-458. (Abstract)  
  Halaj, J., C. B. Halpern, and H. Yi. 2009. Effects of green-tree retention on abundance and guild composition ofcorticolous arthropods. Forest Ecology and Management 258:850-859. (Abstract)  
  Harrod, R. J., and C. B. Halpern. 2009. Effects of experimental burning on individual performance and population structure of two rare plants of northcentral Washington. Restoration Ecology 17:215-225. (Abstract)  
  Urgenson, L., S. Reichard, and C. Halpern. 2009. Community and ecosystem consequences of giant knotweed (Polygonum sachalinense) invasion into riparian forests of western Washington, USA. Biological Conservation 142:1536-1541. (Abstract)  
  Halaj, J., C. B. Halpern, and H. Yi. 2008. Responses of litter-dwelling spiders and carabid beetles to varying levels and  patterns of green-tree retention. Forest Ecology and Management 255:887-900.(Abstract)  
  Jones, C. C., C. B. Halpern, and J. Niederer. 2008. Plant succession on gopher mounds in western Cascade meadows: consequences for species diversity and heterogeneity. American Midland Naturalist 159:275-286. (Abstract)  
  Nelson, C. R., C. B. Halpern, and J. K. Agee. 2008. Thinning and burning result in limited invasion of nonnative plants but neutral effects on natives. Ecological Applications 18:762-770. (pdf)  
  Six, L. J., and C. B. Halpern. 2008. Substrate effects on distribution, biomass allocation and morphology of forest understory plants. Botany 86:1133-1142. (Abstract)  
  Gitzen, R. A., S. D. West, C. C. Maguire, T. Manning, and C. B. Halpern. 2007. Response of small mammals to varying amounts and patterns of green-tree retention. Forest Ecology and Management 251:142-155.(Abstract)  
  Haugo, R. D., and C. B. Halpern. 2007. Vegetation responses to conifer encroachment in a dry, montane meadow: a chronosequence approach. Canadian Journal of Botany 85:285-298. (pdf)  
  Heithecker, T. D., and C. B. Halpern. 2007. Edge-related gradients in microclimate in forest aggregates following structural retention harvests in western Washington. Forest Ecology and Management 248:163-173. (Abstract)  
  Lang, N. L., and C. B. Halpern. 2007. The soil seed bank of a montane meadow: consequences of conifer encroachment and implications for restoration. Canadian Journal of Botany 85:557-569. (pdf)  
  Maguire, D. A., C. B. Halpern, and D. L. Phillips. 2007. Changes in forest structure following variable-retention harvests: target and non-target effects. Forest Ecology and Management 242:708-726.(Abstract)  
  Nelson, C. R., C. B. Halpern, and J. A. Antos. 2007. Variation in responses of late-seral herbs to disturbance and environmental stress. Ecology 88:2880-2890. (pdf)  
 

Dovciak, M., C. B. Halpern, J. F. Saracco, S. A. Evans, and D. A. Liguori. 2006. Persistence of ground-layer bryophytes  in a structural-retention experiment: initial effects of level and pattern of retention. Canadian Journal of Forest Research 36:3039-3052. (pdf)

 
  Heithecker, T. D., and C. B. Halpern. 2006. Variation in microclimate associated with dispersed-retention harvests in coniferous forests of western Washington. Forest Ecology and Management 226:60-71. (pdf)  
 

Lutz, J. A., and C. B. Halpern. 2006. Tree mortality during early forest development: a long-term study of rates, causes, and consequences. Ecological Monographs 76:257-275. (pdf)

 
  Maguire, D. A., D. B. Mainwaring, and C. B. Halpern. 2006. Stand dynamics after variable-retention harvesting in mature   Douglas-fir forests of western North America. Allgemeine Forst und Jagdzeitung 177:120-131. (pdf)  
  Halpern, C. B., D. McKenzie, S. A. Evans, and D. A. Maguire. 2005. Initial responses of forest understories to varying levels and patterns of green-tree retention. Ecological Applications 15:175-195. (pdf)  
  Harrod, R. J., and C. B. Halpern. 2005. Effects of experimental shading and removal of competitors on vegetative and reproductive performance of Trifolium thompsonii Morton, a rare endemic herb of eastern Washington. Natural Areas Journal 25:156-164.
 
  Harrod, R. J., and C. B. Halpern. 2005. The seed ecology of Iliamna longisepala (Torr.) Wiggins, an east Cascade endemic Natural Areas Journal 25:246-256.  
  Nelson, C. R., and C. B. Halpern. 2005. Edge-related responses of understory species to aggregated retention harvest in the Pacific Northwest. Ecological Applications 15:196-209. (pdf)  
  Nelson, C. R., and C. B. Halpern. 2005. Short-term effects of timber harvest and forest edges on ground-layer mosses and liverworts. Canadian Journal of Botany 83:610-620. (pdf)  
  Aubry, K.B., C.B. Halpern, and D.A. Maguire. 2004. Ecological effects of variable-retention harvests in the northwestern United States: the DEMO study. Forest Snow and Landscape Research 78:119-134. (pdf)  
  Antos, J. A., C. B. Halpern, R. E. Miller, K. Cromack, Jr., and M. G. Halaj. 2003. Temporal and spatial changes in soil carbon and nitrogent after clearcutting and burning of an old-growth Douglas-fir forest. USDA Forest Service Research Paper PNW-RP-552. (pdf)  
  Moore, J. R., D. A. Maguire, D. L. Phillips, and C. B. Halpern. 2002. Effects of varying levels and patterns of green-tree retention on amount of harvesting damage. Western Journal of Applied Forestry 17:202–206. (Abstract)  
  Acker, S. A., C. B. Halpern, M. E. Harmon, and C. T. Dyrness. 2002. Trends in bole biomass accumulation, net primary production, and tree mortality in Pseudotsuga menziesii forests of contrasting age. Tree Physiology 22:213-217. (Abstract)  
  Halpern, C.B., and D. McKenzie. 2001. Disturbance and post-harvest ground conditions in a structural retention experiment Forest Ecology and Management 154:215-225. (Abstract)  
  Lezberg, A. L., C. B. Halpern, and J. A. Antos. 2001. Clonal development of Maianthemum dilatatum in forests of differing age and structure. Canadian Journal of Botany 79:1028-1038. (pdf)  
  Thomas, S. C., D. A. Liguori, and C. B. Halpern. 2001. Corticolous bryophytes in managed Douglas-fir forests: habitat differentiation and responses to thinning and fertilization. Canadian Journal of Botany 79:886-896. (pdf)  
  Beach, E. W., and C. B. Halpern. 2001. Controls on conifer regeneration in managed riparian forests: effects of seed source, substrate, and vegetation. Canadian Journal of Forest Research 31:471-482. (pdf)  
 
McKenzie, D., C. B. Halpern, and C. R. Nelson. 2000. Overstory influences on herb and shrub communities in mature forests of western Washington, USA. Canadian Journal of Forest Research 30:1655-1666. (pdf)
 
  Aubry, K. B., M. P. Amaranthus, C. B. Halpern, J. D. White, B. L. Woodard, C. E. Peterson, C. A. Lagoudakis, and A. J. Horton. 1999. Experimental design of the DEMO project: A study of varying levels and patterns of green-tree retention in harvest units in Oregon and Washington. Northwest Science 73 (Special Issue):12-26. (pdf)  
  Franklin, J. F., and C. B. Halpern. 1999. Pacific Northwest Forests. Pages 123-159 in M. G. Barbour and W. D. Billings (editors). North American Terrestrial Vegetation (second edition). Cambridge University Press, New York.  
  Halpern, C. B., and M. G. Raphael, editors. 1999. Special issue on retention harvests in Northwestern forest ecosystems:  The Demonstration of Ecosystem Management Options (DEMO) Study. Northwest Science 73 (Special Issue):1-2.
  Halpern, C. B., S. A. Evans, C. R. Nelson, D. McKenzie, D. A. Liguori, D. E. Hibbs, and M. G. Halaj. 1999. Response of forest vegetation to varying levels and patterns of green-tree retention: An overview of a long-term experiment. Northwest Science 73 (Special Issue):27-44. (pdf)
  Halpern, C. B., S. A. Evans, and S. Nielson. 1999. Soil seed banks in young, closed-canopy forests of the Olympic Peninsula, Washington: Potential contributions to understory reinitiation. Canadian Journal of Botany 77:922-935. (pdf)
  Lezberg, A. L., J. A. Antos, and C. B. Halpern. 1999. Below-ground traits of herbaceous species in young, coniferous forests of the Olympic Peninsula, Washington. Canadian Journal of Botany 77: 936-943. (pdf)
  McKenzie, D., and C. B. Halpern. 1999. Modeling the distributions of shrub species in Pacific Northwest forests. Forest Ecology and Management 114:293-307.(Abstract)
  Thomas, S. C., C. B. Halpern, D. A. Falk, D. A. Liguori, and K. A. Austin. 1999. Plant diversity in managed forests: understory responses to thinning and fertilization. Ecological Applications 9:864-879. (pdf)
  Miller, E. A., and C. B. Halpern. 1998. Effects of environment and grazing disturbance on tree establishment in meadows of the western Cascade Range, Oregon, USA. Journal of Vegetation Science 9:265-282.(pdf)
  Antos, J. A., and C. B. Halpern. 1997. Root system differences among species: implications for early successional changes in forests of western Oregon. American Midland Naturalist 138:97-108.
  Halpern, C. B., J. A. Antos, M. A. Geyer, and A. M. Olson. 1997. Species replacement during early secondary succession: the abrupt decline of winter annual. Ecology 78:621-631. (pdf)
  Halpern, C. B., E. A. Miller, and M. A. Geyer. 1996. Equations for predicting above-ground biomass of plant species in early successional forests of the western Cascade Range, Oregon. Northwest Science 70:306-320. (pdf)
  Halpern, C. B., and T. A. Spies. 1995. Plant species diversity in natural and managed forests of the Pacific Northwest. Ecological Applications 5:913-934. (pdf)
  Busing, R. T., C. B. Halpern, and T. A. Spies. 1995. Ecology of Pacific yew (Taxus brevifolia) in western Oregon and Washington. Conservation Biology 9:1199-1207. (Abstract)  
  Urban, D. L., M. E. Harmon, and C. B. Halpern. 1993. Potential response of Pacific Northwestern forests to climatic change, effects of stand age and initial composition. Climatic Change 23: 247-266.  
  Halpern, C. B., J. A. Antos, K. Cromack, Jr., and A. M. Olson. 1992. Species' interactions and plant diversity during secondary succession. The Northwest Environmental Journal 8:203-205.  
  Halpern, C. B., J. F. Franklin, and A. McKee. 1992. Changes in plant species diversity after harvest of Douglas-fir forests. The Northwest Environmental Journal 8:205-207.  
  Taylor, A. H., and C. B. Halpern. 1991. The structure and dynamics of red fir (Abies magnifica) forests, southern Cascade  Range, USA. Journal of Vegetation Science 2:189-200. (Abstract)  
  Halpern, C. B., and J. F. Franklin. 1990. Physiognomic development of Pseudotsuga forests in relation to initial structure and disturbance intensity. Journal of Vegetation Science 1:475-482. (Abstract)  
  Halpern, C. B., P. M. Frenzen, J. E. Means, and J. F. Franklin. 1990. Plant succession in areas of scorched and blown-down forest after the 1980 eruption of Mount St. Helens, Washington. Journal of Vegetation Science 1:181-194. (Abstract)  
  Halpern, C. B. 1989. Early successional patterns of forest species: interactions of life history traits and disturbance. Ecology 70:704-720. (pdf)  
  Halpern, C. B. 1988. Early successional pathways and the resistance and resilience of forest communities. Ecology 69:1703-1715. (pdf)  
  Halpern, C. B. 1986. Montane meadow plant associations of Sequoia National Park, California. Madroño 33:1-23.  
  Halpern, C. B., B. G. Smith, and J. F. Franklin. 1984. Composition, structure, and distribution of the ecosystems of the Three Sisters Biosphere Reserve/Wilderness Area. A Final Report to the United States Department of Agriculture. (pdf)  
  Halpern, C. B., and M. E. Harmon. 1983. Early plant succession on the Muddy River Mudflow, Mount St. Helens, Washington. American Midland Naturalist 110:97-106.