Jonathan completed his Master’s of Science in Geology at the University of Oregon in 2010. His interests include Cenozoic mammals, dental microwear, and vertebrate paleontology. He advanced to PhD candidacy in March 2012 and graduated in 2016. His PhD research focused on the paleoecology of mammalian faunas of the Oligo-Miocene Renova Formation in southwest Montana in comparison with patterns from the John Day Fossil Beds of Oregon and the Arikaree Group of the Central Great Plains. His work has been funded by the GSA, the American Philosophical Society, The Paleontological Society, Sigma Xi, and others. Check out his page.